Evolution in SNAPDRAGONS

Linaria - Linaria bipunctata - Taxonomy

Linaria bipunctata (L.) Chaz., Suppl. Dict. Jard. 2: 39 (1790).

Somewhat glaucous annual or biennial, glandular-puberulent in inflorescence, glandular­puberulent to subglabrous below; fertile stems 5-20 (-40), 14-32 cm, more or less erect to ascending, slender, diffusely branched above; sterile stems 0-12. Leaves of fertile stems flat or somewhat revolute, in whorls of 4-5 below, alternate above; leaves of sterile stems similar to those of fertile stems, mostly in whorls of 3-4. Inflorescence with 3-8 flowers, somewhat dense in flower, very lax in fruit. Bracts 1-3 mm, linear, subobtuse. Pedicels erect. Calyx-lobes somewhat unequal, linear-lanceolate to oblong, subobtuse; adaxial lobe 1.8-3 x 0.3-0.4 mm in flower and fruit; abaxial lobes 0.8-2.5 x 0.4-0.7 mm in flower and fruit. Corolla (9-) 11-16 mm, yellow with deeper yellow palate; tube 1.5-2.5 mm broad; adaxial lip sinus 1-1.5 mm; abaxial lip sinus 0.7-1.4 mm; spur 4-6.5 mm, 0.6-1 mm broad, straight or somewhat curved, much shorter than rest of corolla, often reddish. Capsule 1.5-3 (-3.5) mm, oblong-ovoid, glandular­puberulent above. Seeds tuberculate and papillate; hilum narrow; periclinal wall of testa-cells tabular, convex or papillate, densely verrucate, the anticlinal wall of testa-cells angular with ridge over junction. Modified from Sutton (1988).

N & C Portugal, C & W Spain. Hs Lu.

Type: SPAIN: “10 bipunctatum” Loefling 488 (syn.? (cf. Valdés, 1970a: 241-242; Viano, 1978b: 217) LINN-767.17!).


a. Linaria bipunctata subsp. bipunctata.


Fertile stems subglabrous or sparsely glandular-pubescent below, branches diverging at a narrow angle throughout the length of stems. Leaves of fertile stems 6-20 x 0.3-0.5 (-1) mm, linear or subulate, evenly spaced along stems. Pedicels 1-3 mm. Seeds (0.5-) 0.7-0.8 mm, reniform-orbicular or somewhat trigonous, usually compressed with an encircling marginal ridge, blackish-grey; marginal ridge up to 0.05 mm broad, somewhat thin, wing-like; tubercles discrete, the distance between tubercles 1-2 x width of tubercles.


E Portugal, C & W Spain. Hs Lu.


b. Linaria bipunctata subsp. glutinosa (Hoffmanns. & Link) D.A. Sutton, stat. nov.


Basionym: Linaria glutinosa Hoffmanns. & Link, Fl. Port. 1(8): 237, tab. 39 (1811), excl. omn. syn.


Fertile stems densely glandular-pubescent throughout, the lowermost 4-6 cm unbranched, divaricately and often diffusely branched above. Leaves of fertile stems 2-9 x 0.6-2.5 mm, linear-lanceolate, widely spaced above, very crowded and caducous below. Pedicels 0.5-1.2 (-1.6) mm, crowded below, widely spaced above. Seeds 0.4-0.6 mm, reniform to reniform­tetrahedral, usually without marginal ridges, black, shiny; tubercles more or less contiguous, often in curved rows.


W Portugal, NW Spain? Hs? Lu.

Types: PORTUGAL: sur le bord de la mer près de Setuval [Setubal, Estremadura] (syn. B?); sur les rochers dan s les contrées maritimes près d O-Porto [Porto, Douro Litoral], iv-vi (syn. B?).

Linaria species
Linaria aeruginea
Linaria albifrons
Linaria algarviana
Linaria alpina
Linaria altaica
Linaria amethystea
Linaria amoi
Linaria angustissima
Linaria anticaria
Linaria antilibanotica
Linaria arenaria
Linaria armeniaca
Linaria arvensis
Linaria atlantica
Linaria badalii
Linaria baligaliensis
Linaria bamianica
Linaria biebersteinii
Linaria bipartita
Linaria bipunctata
Linaria bordiana
Linaria brachyceras
Linaria bubanii
Linaria bungei
Linaria buriatica
Linaria caesia
Linaria capraria
Linaria cavanillesii
Linaria chalepensis
Linaria clementei
Linaria confertiflora
Linaria corifolia
Linaria cossonii
Linaria coutinhoi
Linaria cretacea
Linaria creticola
Linaria dalmatica
Linaria damascena
Linaria debilis
Linaria decipiens
Linaria depauperata
Linaria diffusa
Linaria dissita
Linaria elegans
Linaria elymaitica
Linaria fallax
Linaria fastigiata
Linaria faucicola
Linaria ficalhoana
Linaria filicaulis
Linaria flava
Linaria genistifolia
Linaria gharbensis
Linaria glacialis
Linaria glauca
Linaria grandiflora
Linaria griffithsii
Linaria haelava
Linaria hellenica
Linaria hepatica
Linaria hirta
Linaria huteri
Linaria iconia
Linaria incarnata
Linaria incompleta
Linaria intricata
Linaria japonica
Linaria joppensis
Linaria kokanica
Linaria kulabensis
Linaria kurdica
Linaria latifolia
Linaria laxiflora
Linaria leptoceras
Linaria lilacina
Linaria lineolata
Linaria loeselii
Linaria macrophylla
Linaria macroura
Linaria maroccana
Linaria melampyroides
Linaria meyeri
Linaria michauxii
Linaria micrantha
Linaria microsepala
Linaria miller
Linaria multicaulis
Linaria munbyana
Linaria nivea
Linaria nurensis
Linaria oblongifolia
Linaria odora
Linaria oligantha
Linaria onubensis
Linaria paradoxa
Linaria parviracemosa
Linaria pedicellata
Linaria pedunculata
Linaria pelisseriana
Linaria peloponnesiaca
Linaria peltieri
Linaria pinifolia
Linaria platycalyx
Linaria polygalifolia
Linaria popovii
Linaria propinqua
Linaria pseudolaxiflora
Linaria pseudoviscosa
Linaria purpurea
Linaria pyramidalis
Linaria ramosa
Linaria reflexa
Linaria remotiflora
Linaria repens
Linaria ricardoi
Linaria riffea
Linaria rubioides
Linaria salzmannii
Linaria satureioides
Linaria saxatilis
Linaria schelkownikowii
Linaria schirvanica
Linaria sessilis
Linaria simplex
Linaria spartea
Linaria striatella
Linaria supina
Linaria tarhunensis
Linaria tenuis
Linaria thibetica
Linaria thymifolia
Linaria tingitana
Linaria tonzigii
Linaria transiliensis
Linaria triornithophora
Linaria triphylla
Linaria tristis
Linaria tuberculata
Linaria tursica
Linaria unaiensis
Linaria venosa
Linaria ventricosa
Linaria veratrifolia
Linaria verticillata
Linaria virgata
Linaria viscosa
Linaria vulgaris
Linaria warionis
Linaria weilleri
Linaria yunnanensis

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