Evolution in SNAPDRAGONS

 
Linaria - Linaria genistifolia - Taxonomy

Linaria genistifolia (L.) Miller, Gard. Dict. ed. 8: no. 14 (1768). 

Glaucous perennial, glabrous or occasionally glandular-pubescent on calyces and capsules, reproducing vegetatively by rhizomes; fertile stems 1-numerous, 20-120 cm, erect or ascending, often branched above; sterile stems usually absent. Leaves of fertile stems acute, alternate. Bracts 3-6 mm, lanceolate, acute. Pedicels erect, 0.5-3 (-4) mm in flower, 3-5 (-8) mm in fruit. Calyx-lobes 3-5 (-6.5) x 0.8-1.2 (-2) mm, lanceolate to ovate, acute. Corolla usually yellow; adaxial lip sinus 0.8-2 mm; abaxial lip sinus 0.7-1.5 mm; spur straight or somewhat curved. Capsule 2.5-6 x 2.5-6.5 mm, subglobose, truncate or emarginate. Seeds (0.7-) 0.9-1.4 mm, crescenti-trigonous or tetrahedral, black or blackish-brown; longitudinal marginal ridges 0.1-0.15 mm high, obtuse to acute, the intervening faces with anastomosed ridges or ruminate-alveolate, sometimes with discrete tubercles; periclinal wall of testa-cells tabular or tabular-convex, densely rugulate to reticulate, that of cells from interstices and ridge-bases usually with median papilla, the papilla hemispherical or conical, rounded to subacute. Modified from Sutton (1988).

C & E Europe to W Asia, widely naturalized elsewhere. Al An Au Bu Cz Gr Hu It Ju Po Rm Rs(Ec Ee Ek Ew Ka) Tu [Br Cn Ga Ge Us].

Type: [USSR]: "genistifolium" (Siberia) [Gerber? s.n.] (lecto. (Davis 1978a: 6) LINN-767.40!).

 

a. Linaria genistifolia subsp. genistifolia.

 

Fertile stems erect or ascending. Leaves of fertile stems 24-50 (-85) x 3-10 (-35) mm, the length (2-) 3.5-8.5 x breadth, linear-lanceolate, 1- to 3-veined, cuneate or attenuate, the lowermost sometimes ovate, 5-veined, flat, semi-amplexicaul. Inflorescence with (5-) 10-50 flowers, usually lax in flower and fruit. Calyx-lobes 3-5 (-6.5) x 0.7-2 mm, glabrous. Corolla (13-) 16-23 mm, bright yellow; tube 3-4.5 mm broad; spur (5-) 7-10 rnm, 1.3-1.7 (-1.8) mm broad at base, shorter than or subequalling rest of corolla. Capsule glabrous.

 

2n = 12 (Heitz, 1926; Valdés, 1970b, Van Loon & Van Setten, 1982: 592).

Throughout most of the range of the species. Al An Au Bu Cz Hu It Ju Po Rm Rs(Ec Ee Ek Ew Ka) Tu [Br Cn Ga Ge Us].

 

b. Linaria genistifolia subsp. artvinensis P.H. Davis, Notes Roy. Bot. Gard. Edinb. 36(1): 7 (1978).

 

Fertile sterns erect. Leaves of fertile stems 10-35 x 5-10 mm, the length 2.5-5 x breadth, ovate to ovate-elliptic, thick-textured, 3-veined, flat, semi-amplexicaul. Inflorescence with numerous flowers, lax in flower and fruit, often secund. Calyx-lobes 2.5-3 x 1.3-1.8 mm, lanceolate to ovate, acute, glabrous. Corolla 13-16 mm, yellow with red veins; tube 2.5-3.5 mm broad; spur 7-8.5 mm, 1-1.5 mm broad at base, scarcely exceeding rest of corolla. Capsule glabrous.

 

NE Anatolia. An.

Type: TURKEY: (A8) Çoruh; Artvin, 400 m, 28 iv 1960 Stainton 8290 (holo. E!, iso. K!).

 

c. Linaria genistifolia subsp. confertiflora (Boiss.) P.H. Davis, Notes Roy. Bot. Gard. Edinb. 36(1): 6 (1978).

Fertile stems erect. Leaves of fertile stems 18-40 (-75) x 3-10 (-16) mm, the length 4-7 x breadth, linear-lanceolate, flat, attenuate. Inflorescence with numerous flowers, dense in flower, dense or somewhat lax in fruit. Calyx-lobes 4-5.2 x 0.6-1.8 mm, linear-lanceolate, acuminate, sparsely glandular-puberulent, -pubescent, -ciliate or rarely glabrous. Corolla 16-19 mm, pale yellow; tube 3.5-4.5 mm broad; spur 7-8 mm, 1-1.3 mm broad at base, somewhat shorter than rest of corolla. Capsule glandular-puberulent or sometimes glabrous.

 

SW & C Anatolia. An.

Type: TURKEY: (C3)Konya; in campis Pisidae et Isauriae, rara, circa Beycheher [Beyşehir], vi 1845 Heldreich s.n. (lectotype selected here G!, iso. BM! E! K!).

 

d. Linaria genistifolia subsp. euxina (Velen.) D.A. Sutton, comb. et stat. nov.

 

Basionym: Linaria euxina Velen., Abh. Boehm. Ges. Wiss. 7(1): 34 (1886).

 

Fertile stems ascending to suberect. Leaves of fertile stems 9-24 (-65) x 0.5-4 (-6) mm, the length (4.5-) 8-20 (-30) x breadth, linear or filiform to linear-lanceolate, flat or involute, attenuate. Inflorescence with 2-25 (-40) flowers, somewhat dense to lax in flower, lax in fruit. Calyx-lobes 2-4.5 x 0.5-1.5 mm, linear or linear-lanceolate, acuminate, glabrous. Corolla 16-18 mm, bright yellow; tube 2.5-3 mm broad; spur 6-8 mm, 0.8-1.2 mm broad at base, shorter than rest of corolla. Capsule glabrous.

 

SE Europe, Anatolia. An Bu Gr Rm Rs(Ek Ka) Tu.

Type: BULGARIA: auf den Sandfluren am Meere bei Varna, viii 1885 Velenovský s.n. (not located).

 

e. Linaria genistifolia subsp. polyclada (Fenzl) P.H. Davis, Notes Roy .Bot. Gard. Edinb. 36(1): 7 (1978).

 

Fertile stems erect. Leaves of fertile stems (l0-) 20-42 x (0.6) 1-1.8 mm, the length (l0-)

20-40 x breadth, linear or filiform, the margins flat or somewhat involute, attenuate. Inflorescence with 6-12 flowers, lax in flower and fruit. Calyx-lobes 2-3.5 x 0.8-1.1 mm, lanceolate, acute, glabrous. Corolla 11-12 mm, pale yellow; tube 2.7-3 mm broad; spur 3.5-4.5 mm, 0.6-1.5 mm broad at base, much shorter than rest of corolla. Capsule glabrous.

 

S & C Anatolia. An.

Type: TURKEY: (C5) Içel; Ciliciae Bulgardagh [Bulgar Dağ], inter segetes prope pagum Gulek [Güllek], 1235 m, 1836 Kotschy 399 (iso. E! G!).

 

f. Linaria genistifolia subsp. praealta (Boiss. & Bal.) P.H. Davis, Notes Roy. Bot. Gard. Edinb. 36(1): 7 (1978).

 

Fertile stems erect. Leaves of fertile stems 20-40 x (3.5-) 6-17 mm, the length 2-5 x breadth, lanceolate to ovate, flat, semi-amplexicaul. Inflorescence with 8-50 flowers, lax in flower and fruit. Calyx-lobes 3.5-5 x 1-2 mm, lanceolate to ovate, acute, glabrous. Corolla 18-21 mm, pale yellow with prominent reddish-brown veins, the adaxial lip often tinged with red; tube 3.5-4.5 mm broad; spur 6-9 mm, 1.3-2 mm broad at base, shorter than rest of corolla. Capsule glabrous or occasionally glandular-pubescent.

SC to SE Anatolia (Taurus, Amanus). An.

Type: TURKEY: (C5) Içel; in fauce Guzel Deré [Güzel Dere] supra pagum Kechlick [Keklik] non procul a Mersina in Cilicia littorali, vi 1855 Balansa s.n. (holo. G!).

Linaria species
Linaria aeruginea
Linaria albifrons
Linaria algarviana
Linaria alpina
Linaria altaica
Linaria amethystea
Linaria amoi
Linaria angustissima
Linaria anticaria
Linaria antilibanotica
Linaria arenaria
Linaria armeniaca
Linaria arvensis
Linaria atlantica
Linaria badalii
Linaria baligaliensis
Linaria bamianica
Linaria biebersteinii
Linaria bipartita
Linaria bipunctata
Linaria bordiana
Linaria brachyceras
Linaria bubanii
Linaria bungei
Linaria buriatica
Linaria caesia
Linaria capraria
Linaria cavanillesii
Linaria chalepensis
Linaria clementei
Linaria confertiflora
Linaria corifolia
Linaria cossonii
Linaria coutinhoi
Linaria cretacea
Linaria creticola
Linaria dalmatica
Linaria damascena
Linaria debilis
Linaria decipiens
Linaria depauperata
Linaria diffusa
Linaria dissita
Linaria elegans
Linaria elymaitica
Linaria fallax
Linaria fastigiata
Linaria faucicola
Linaria ficalhoana
Linaria filicaulis
Linaria flava
Linaria genistifolia
Linaria gharbensis
Linaria glacialis
Linaria glauca
Linaria grandiflora
Linaria griffithsii
Linaria haelava
Linaria hellenica
Linaria hepatica
Linaria hirta
Linaria huteri
Linaria iconia
Linaria incarnata
Linaria incompleta
Linaria intricata
Linaria japonica
Linaria joppensis
Linaria kokanica
Linaria kulabensis
Linaria kurdica
Linaria latifolia
Linaria laxiflora
Linaria leptoceras
Linaria lilacina
Linaria lineolata
Linaria loeselii
Linaria macrophylla
Linaria macroura
Linaria maroccana
Linaria melampyroides
Linaria meyeri
Linaria michauxii
Linaria micrantha
Linaria microsepala
Linaria miller
Linaria multicaulis
Linaria munbyana
Linaria nivea
Linaria nurensis
Linaria oblongifolia
Linaria odora
Linaria oligantha
Linaria onubensis
Linaria paradoxa
Linaria parviracemosa
Linaria pedicellata
Linaria pedunculata
Linaria pelisseriana
Linaria peloponnesiaca
Linaria peltieri
Linaria pinifolia
Linaria platycalyx
Linaria polygalifolia
Linaria popovii
Linaria propinqua
Linaria pseudolaxiflora
Linaria pseudoviscosa
Linaria purpurea
Linaria pyramidalis
Linaria ramosa
Linaria reflexa
Linaria remotiflora
Linaria repens
Linaria ricardoi
Linaria riffea
Linaria rubioides
Linaria salzmannii
Linaria satureioides
Linaria saxatilis
Linaria schelkownikowii
Linaria schirvanica
Linaria sessilis
Linaria simplex
Linaria spartea
Linaria striatella
Linaria supina
Linaria tarhunensis
Linaria tenuis
Linaria thibetica
Linaria thymifolia
Linaria tingitana
Linaria tonzigii
Linaria transiliensis
Linaria triornithophora
Linaria triphylla
Linaria tristis
Linaria tuberculata
Linaria tursica
Linaria unaiensis
Linaria venosa
Linaria ventricosa
Linaria veratrifolia
Linaria verticillata
Linaria virgata
Linaria viscosa
Linaria vulgaris
Linaria warionis
Linaria weilleri
Linaria yunnanensis

 
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