Evolution in SNAPDRAGONS

Linaria - Linaria macrophylla - Taxonomy

Linaria macrophylla Kuprian., Acta Inst. Bot. Acad. Sci. URSS ser. 1 9: 63, tab. 2(4) (1950).

Glabrous, somewhat glaucous perennial; fertile stems (1-) 34, 20-40 cm, suberect, robust, simple or sparingly branched above, the branches usually verticillate; sterile stems apparently absent. Leaves (5-) 8-20 x (4.5-) 8-20 mm, the length 0.8-1.1 x breadth, suborbicular or reniform-orbicular, obtuse to abruptly acuminate, flat, amplexicaul or subauriculate, thick-textured, mostly 3- or 4-verticillate below, the uppermost opposite or alternate. Inflorescence with 3-18 flowers, more or less dense in flower and fruit. Bracts 1-2 mm, linear, acute. Pedicels 1-3 mm in flower and fruit, erect. Calyx-lobes 2-2.5 x 0.6-1 mm, linear-oblong to lanceolate, acute. Corolla 8.5-10 mm, yellow; tube 2.5-3 mm broad; adaxial lip sinus 1.5-1.8 mm; abaxial lip sinus 1.2-1.5 mm; spur 3.5-4 mm, 0.6-1 mm broad at base, more or less straight, much shorter than rest of corolla. Capsule 2-3 x 2-3 mm, globose, glabrous. Seeds 1.1-1.5 mm, crescenti-trigonous or oblong-reniform, strongly compressed, black; longitudinal marginal ridges 2-3, 0.1-0.2 mm high, somewhat wing-like, the intervening faces finely tuberculate or with sinuate, anastomosed, ridges; periclinal wall of testa cells, tabular, rugulate, not papillate. Modified from Sutton (1988).

Soviet Central Asia. Rs(Ca)

Type: USSR: Aktyubinsk; (Kazakhstan) distr. Temir, mons cretacea Astau-saldy, 19 vi 1904 Dubiansky 850 (holo. LE, photo. BM!).

Described from calcareous substrate on the banks of the river Emba in western Kazakhstan, L. macrophylla resembles L. cretacea except for the taller stems, broader, more obtuse, amplexicaul leaves and smaller flowers.


Linaria species
Linaria aeruginea
Linaria albifrons
Linaria algarviana
Linaria alpina
Linaria altaica
Linaria amethystea
Linaria amoi
Linaria angustissima
Linaria anticaria
Linaria antilibanotica
Linaria arenaria
Linaria armeniaca
Linaria arvensis
Linaria atlantica
Linaria badalii
Linaria baligaliensis
Linaria bamianica
Linaria biebersteinii
Linaria bipartita
Linaria bipunctata
Linaria bordiana
Linaria brachyceras
Linaria bubanii
Linaria bungei
Linaria buriatica
Linaria caesia
Linaria capraria
Linaria cavanillesii
Linaria chalepensis
Linaria clementei
Linaria confertiflora
Linaria corifolia
Linaria cossonii
Linaria coutinhoi
Linaria cretacea
Linaria creticola
Linaria dalmatica
Linaria damascena
Linaria debilis
Linaria decipiens
Linaria depauperata
Linaria diffusa
Linaria dissita
Linaria elegans
Linaria elymaitica
Linaria fallax
Linaria fastigiata
Linaria faucicola
Linaria ficalhoana
Linaria filicaulis
Linaria flava
Linaria genistifolia
Linaria gharbensis
Linaria glacialis
Linaria glauca
Linaria grandiflora
Linaria griffithsii
Linaria haelava
Linaria hellenica
Linaria hepatica
Linaria hirta
Linaria huteri
Linaria iconia
Linaria incarnata
Linaria incompleta
Linaria intricata
Linaria japonica
Linaria joppensis
Linaria kokanica
Linaria kulabensis
Linaria kurdica
Linaria latifolia
Linaria laxiflora
Linaria leptoceras
Linaria lilacina
Linaria lineolata
Linaria loeselii
Linaria macrophylla
Linaria macroura
Linaria maroccana
Linaria melampyroides
Linaria meyeri
Linaria michauxii
Linaria micrantha
Linaria microsepala
Linaria miller
Linaria multicaulis
Linaria munbyana
Linaria nivea
Linaria nurensis
Linaria oblongifolia
Linaria odora
Linaria oligantha
Linaria onubensis
Linaria paradoxa
Linaria parviracemosa
Linaria pedicellata
Linaria pedunculata
Linaria pelisseriana
Linaria peloponnesiaca
Linaria peltieri
Linaria pinifolia
Linaria platycalyx
Linaria polygalifolia
Linaria popovii
Linaria propinqua
Linaria pseudolaxiflora
Linaria pseudoviscosa
Linaria purpurea
Linaria pyramidalis
Linaria ramosa
Linaria reflexa
Linaria remotiflora
Linaria repens
Linaria ricardoi
Linaria riffea
Linaria rubioides
Linaria salzmannii
Linaria satureioides
Linaria saxatilis
Linaria schelkownikowii
Linaria schirvanica
Linaria sessilis
Linaria simplex
Linaria spartea
Linaria striatella
Linaria supina
Linaria tarhunensis
Linaria tenuis
Linaria thibetica
Linaria thymifolia
Linaria tingitana
Linaria tonzigii
Linaria transiliensis
Linaria triornithophora
Linaria triphylla
Linaria tristis
Linaria tuberculata
Linaria tursica
Linaria unaiensis
Linaria venosa
Linaria ventricosa
Linaria veratrifolia
Linaria verticillata
Linaria virgata
Linaria viscosa
Linaria vulgaris
Linaria warionis
Linaria weilleri
Linaria yunnanensis

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