Evolution in SNAPDRAGONS

 
Linaria - Linaria pyramidalis - Taxonomy

Linaria pyramidalis (Vent.) F.G. Dietr., Nachtr. Vollsr. Lexic. 4: 417 (1818).

Perennial, glabrous or glabrescent except in inflorescence; fertile stems usually solitary, 30-85 cm, erect or suberect, simple or rarely branched above, stout, densely leafy; sterile stems not seen. Leaves acute or subacute, flat, 3- to 5-veined, alternate. Inflorescence with 14-80 flowers, very dense in flower and fruit. Bracts 5-15 mm, lanceolate to deltate-ovate, acute. Pedicels 1-3 mm in flower and fruit, erect. Calyx-lobes unequal, ovate to oblong-elliptic, obtuse or truncate, the margins not scarious. Corolla bright yellow, glabrous externally; tube 6-11.5 mm broad at mouth; spur straight, shorter than rest of corolla. Capsule 6-8 x 4-6 mm, oblong-globose, glabrous. Seeds reniform-orbicular, grey to black; wing pale grey; disc reniform, tuberculate; tubercles conical, occasionally laterally compressed or anastomosed to form short, radiating ridges, obtuse to acute. Modified from Sutton (1988).

SW Asia. An Iq Ir Rs(Ck Ka).

Type: [TURKEY/USSR]: "Linaria Armenia lilii Persici foliis florum spica densissima et pyramidata" (Linaria orientalis lilii persici foliis, florum spica densissima et pyramidata) Tournefort (holo. P-Tourn. 967, photo. E!, microfiche IDC-6208/48/12!, type collection BM!).

 

a. Linaria pyramidalis subsp. pyramidalis.

Idorescence subglabrous to sparsely glandular-villous. Leaves 40-85 x 5-15 (-20) mm, the length (3.5-) 5-8 (-10) x breadth, lanceolate, attenuate to cuneate, green or somewhat glaucous. Calyx-lobes obtuse or truncate, subglabrous to somewhat sparsely glandular-villous with brown hairs; adaxial lobe 6.5-8 (-8.5) x 2-4 mm; lateral lobes 5.5-7 (-8) x (2.5-) 3-6 mm. Corolla 28-33 (-37) mm; adaxial lip sinus 2-2.5 mm; abaxial lip sinus 1.5-2 mm; spur 10-14 mm, 2.5-4 mm broad at base. Seeds 1.5-2.6 mm; wing 0.4-0.6 mm broad.

SW Asia (E Anatolia, NW Iran & N Iraq). An Iq Ir.

 

b. Linaria pyramidalis subsp. lenkoranica (Kuprian) D.A. Sutton, comb. et stat. nov.

Basionym: Linaria lenkoranica Kuprian., Acta Inst. Bot. Acad. Sci. URSS ser. 1 9: 67, tab. l(2) (1950).

 

Inflorescence lanate. Leaves 23-50 x (8-) 12-30 mm, the length 2.5-4 x breadth, broadly ovate, cuneate to semi-amplexicaul, very glaucous. Calyx-lobes white lanate, obtuse to acute; adaxial lobe 6.5-8 x 2-2.5 mm; lateral lobes 5-7 x 3-4.5 mm. Corolla 33-40 mm; adaxial lip sinus 1-2.5 mm; abaxial lip sinus 1.5-3; spur 10-18 mm, 1.5-2 mm broad at base. Capsule eglandular-villous; Seeds 2.5-3 mm; wing 0.4-0.7 mm broad.

 

Russia (Transcaucasus). Rs(Ka).

Type: USSR: Azerbaydzhan; distr. Lenkoran, prope pagum Geladaza, decliv. septentr., 13 vii 1931 Matveeva s.n. (holo. LE, photo. BM! E! K!).

 

c. Linaria pyramidalis subsp. kopetdaghensis (Kuprian.) D.A. Sutton, comb. et stat. nov.

Basionym: Linaria kopetdaghensis Kuprian., Acta Inst. Bot. Acad. Sci. URSS ser. 1 9: 66, tab. l(1) (1950).

 

Inflorescence glandular-villous. Leaves 30-50 (-60) x 5-10 (-21) mm, the length 4.5-8 x breadth, linear-lanceolate or broadly lanceolate below, usually attenuate, green. Calyx-lobes glandular-villous with brown hairs; adaxial lobe 8.5-9 (-10) x 2-2.7 mm; lateral lobes 6.5-8 x 3-5 mm. Corolla 36-45 mm; adaxial lip sinus 3.8-4.5 mm; abaxial lip sinus 2.3-2.8 (-3) mm; spur (12-) 14-18 mm, 2.5-4 mm broad at base. Capsule glandular-pubescent. Seeds (2-) 2.7-3 mm; wing 0.7-0.9 mm broad.

Kopet Dagh range of Russia and adjacent parts of Iran. Ir Rs(Ck).

Type: USSR: Turcomania, Kopetdagh, prope Nuchur, 28 vi 1925 Massagetov & Bobrov s.n. (holo. LE).

 

 

Linaria species
Linaria aeruginea
Linaria albifrons
Linaria algarviana
Linaria alpina
Linaria altaica
Linaria amethystea
Linaria amoi
Linaria angustissima
Linaria anticaria
Linaria antilibanotica
Linaria arenaria
Linaria armeniaca
Linaria arvensis
Linaria atlantica
Linaria badalii
Linaria baligaliensis
Linaria bamianica
Linaria biebersteinii
Linaria bipartita
Linaria bipunctata
Linaria bordiana
Linaria brachyceras
Linaria bubanii
Linaria bungei
Linaria buriatica
Linaria caesia
Linaria capraria
Linaria cavanillesii
Linaria chalepensis
Linaria clementei
Linaria confertiflora
Linaria corifolia
Linaria cossonii
Linaria coutinhoi
Linaria cretacea
Linaria creticola
Linaria dalmatica
Linaria damascena
Linaria debilis
Linaria decipiens
Linaria depauperata
Linaria diffusa
Linaria dissita
Linaria elegans
Linaria elymaitica
Linaria fallax
Linaria fastigiata
Linaria faucicola
Linaria ficalhoana
Linaria filicaulis
Linaria flava
Linaria genistifolia
Linaria gharbensis
Linaria glacialis
Linaria glauca
Linaria grandiflora
Linaria griffithsii
Linaria haelava
Linaria hellenica
Linaria hepatica
Linaria hirta
Linaria huteri
Linaria iconia
Linaria incarnata
Linaria incompleta
Linaria intricata
Linaria japonica
Linaria joppensis
Linaria kokanica
Linaria kulabensis
Linaria kurdica
Linaria latifolia
Linaria laxiflora
Linaria leptoceras
Linaria lilacina
Linaria lineolata
Linaria loeselii
Linaria macrophylla
Linaria macroura
Linaria maroccana
Linaria melampyroides
Linaria meyeri
Linaria michauxii
Linaria micrantha
Linaria microsepala
Linaria miller
Linaria multicaulis
Linaria munbyana
Linaria nivea
Linaria nurensis
Linaria oblongifolia
Linaria odora
Linaria oligantha
Linaria onubensis
Linaria paradoxa
Linaria parviracemosa
Linaria pedicellata
Linaria pedunculata
Linaria pelisseriana
Linaria peloponnesiaca
Linaria peltieri
Linaria pinifolia
Linaria platycalyx
Linaria polygalifolia
Linaria popovii
Linaria propinqua
Linaria pseudolaxiflora
Linaria pseudoviscosa
Linaria purpurea
Linaria pyramidalis
Linaria ramosa
Linaria reflexa
Linaria remotiflora
Linaria repens
Linaria ricardoi
Linaria riffea
Linaria rubioides
Linaria salzmannii
Linaria satureioides
Linaria saxatilis
Linaria schelkownikowii
Linaria schirvanica
Linaria sessilis
Linaria simplex
Linaria spartea
Linaria striatella
Linaria supina
Linaria tarhunensis
Linaria tenuis
Linaria thibetica
Linaria thymifolia
Linaria tingitana
Linaria tonzigii
Linaria transiliensis
Linaria triornithophora
Linaria triphylla
Linaria tristis
Linaria tuberculata
Linaria tursica
Linaria unaiensis
Linaria venosa
Linaria ventricosa
Linaria veratrifolia
Linaria verticillata
Linaria virgata
Linaria viscosa
Linaria vulgaris
Linaria warionis
Linaria weilleri
Linaria yunnanensis

 
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