Evolution in SNAPDRAGONS

Linaria - Linaria supina - Taxonomy

Linaria supina (L.) Chaz., Suppl. Dict. Jard. 2: 39 (1790).

Glaucous perennial, sometimes biennial or annual, glabrous except for glandular-pubescent inflorescence; fertile stems often numerous, 5-30 (-40) cm, procumbent to ascending, more rarely suberect, usually simple; sterile stems usually numerous, procumbent to aseending. Leaves of fertile stems attenuate; leaves of sterile sterns similar to those of fertile stems but somewhat smaller and in whorls of 3-6. Inflorescence with 2-12 (-27) flowers, dense in flower, somewhat lax in fruit Bracts 2-11 mm, linear to linear-oblanceolate, acute. Pedicels 0.5-4 (-9.5) mm in flower and fruit, erecto-patent. Calyx-lobes unequal or rarely subequal, subacute to obtuse; adaxial lobe (4-) 4.5-8 x 0.5-1 mm in flower and fruit, linear-oblanceolate to linear­spathulate; abaxial lobes 2.5-6 x 0.8-1.5 mm in flower and fruit, oblanceolate to oblong­ oblanceolate. Corolla pale yellow, occasionally with purp1e or greenish-grey veíns, rarely tinged violet; tube 3-4 mm broad; adaxial lip sinus 2-4 mm; abaxial lip sínus 1.5-3 mm; spur 1.5-2 mm broad at base, slender or rarely stout, somewhat curved, usually exceeding rest of corolla. Capsule 3-6 mm, globose, glabrous or rarely glandular-pubescent. Seeds suborbicular, plano-convex or somewhat concavo-convex; hilum narrow; wing 0.3-0.6 mm broad, thin, smooth, not papillate, black or rarely dark grey; disc somewhat reniform, smooth or tuberculate, not papillate, black, occasionally metallic-shiny, the tubercles sparse and rounded, rarely more dense and acute; anticlinal walls of testa-cells angular with low ridge over junction. Modified from Sutton (1988).

SW Europe. Ga He Hs It Lu (Br Su).

Type: FRANCE: "Linaria pumila supina lutea Bauh. Gallia Narb." (syn. UPS-Herb. Burser XII.29, microfiche IDC-1064/76/12!).


a. Linaria supina subsp. supina.

Perennial or rarely annual; fertile stems numerous or few, ascendíng to suberect, more rarely procumbent. Leaves offertile stems 5-20 (-32) x 0.5-2.5 (-4.5) mm, linear to linear-oblanceolate, acute or subacute, flat or rarely revolute, in whorls of 3-6 below, alternate above, usually equally distributed around stem. Inflorescence with (2-) 6-12 (-27) flowers, dense in flower, somewhat lax in fruit. Corolla (13-) 17-25 (-27) mm; spur (6.5-) 9-13 mm, usually exceeding rest of corolla. Seeds 1.7-2.8 (-3.3) mm; disc smooth or with sparse, rounded tubercles, the tubercle length 0.2-0.8 x width; periclinal wall of testa-cells convex-tabular with prominent margins, smooth or sparsely verruculate.

2n = 12 (Heitz, 1927a: 51; Valdés, 1973: 212).

SW Europe. Ga He Hs It Lu (Br Su).


b. Linaria supina subsp. duriaeana D.A. Sutton, subsp. nov.

A Linaria supina (L.) Chaz. subsp. supina praecipue foliis plerumque alternis secundis, basin caulium versus aggregatis, calcari corollae partem relictam breviore, seminibus tuberculis densis acutis differt.

Herba perennis; caules fertiles numerosi, procumbentes vel interdum ascendentes. Folia caulium fertilium 3-12 (-20) x 0.3-1.2 (-2.2) mm, linearia vel linearo-subulata, apice acuta, margine revoluta, alterna aut basin versus 3-4 in verticillo disposita, plerumque densa secunda. Inflorescentia floribus 1-6 praedita, in statu fiorenti fructiferoque densa. Corolla 15-23 mm longa, calcari 6.5-10 mm longi, corollae parte relicta breviore. Semina 2-2.5 mm longa, disco tuberculis densis acutis, 1-2.5plo longioribus quam latioribus; paries periclinalis cellularum testae tabularis, dense verruculosus.

Perennial; fertile stems numerous, procumbent or sometimes ascending. Leaves of fertile stems 3-12 (-20) x 0.3-1.2 (-2.2) mm, linear or linear-subulate, acute, revolute, alternate or some­times in whorls of 3-4 towards base, usually dense and secund. Inflorescence with 1-6 flowers, dense in flower and fruit. Corolla 15-23 mm; spur 6.5-10 mm, shorter than rest of corolla. Seeds 2-2.5 mm; disc with dense, acute tubercles, the length of tubercle 1-2.5 x width; periclinal wall of testa-cells tabular, densely verruculate.

NW Spain (Oviedo). Hs.

Type: SPAIN: Oviedo; Summis jugis, Pico de Arvas 8 vii 1835 Durieu 259a (holo. BM! iso. K!).


Linaria species
Linaria aeruginea
Linaria albifrons
Linaria algarviana
Linaria alpina
Linaria altaica
Linaria amethystea
Linaria amoi
Linaria angustissima
Linaria anticaria
Linaria antilibanotica
Linaria arenaria
Linaria armeniaca
Linaria arvensis
Linaria atlantica
Linaria badalii
Linaria baligaliensis
Linaria bamianica
Linaria biebersteinii
Linaria bipartita
Linaria bipunctata
Linaria bordiana
Linaria brachyceras
Linaria bubanii
Linaria bungei
Linaria buriatica
Linaria caesia
Linaria capraria
Linaria cavanillesii
Linaria chalepensis
Linaria clementei
Linaria confertiflora
Linaria corifolia
Linaria cossonii
Linaria coutinhoi
Linaria cretacea
Linaria creticola
Linaria dalmatica
Linaria damascena
Linaria debilis
Linaria decipiens
Linaria depauperata
Linaria diffusa
Linaria dissita
Linaria elegans
Linaria elymaitica
Linaria fallax
Linaria fastigiata
Linaria faucicola
Linaria ficalhoana
Linaria filicaulis
Linaria flava
Linaria genistifolia
Linaria gharbensis
Linaria glacialis
Linaria glauca
Linaria grandiflora
Linaria griffithsii
Linaria haelava
Linaria hellenica
Linaria hepatica
Linaria hirta
Linaria huteri
Linaria iconia
Linaria incarnata
Linaria incompleta
Linaria intricata
Linaria japonica
Linaria joppensis
Linaria kokanica
Linaria kulabensis
Linaria kurdica
Linaria latifolia
Linaria laxiflora
Linaria leptoceras
Linaria lilacina
Linaria lineolata
Linaria loeselii
Linaria macrophylla
Linaria macroura
Linaria maroccana
Linaria melampyroides
Linaria meyeri
Linaria michauxii
Linaria micrantha
Linaria microsepala
Linaria miller
Linaria multicaulis
Linaria munbyana
Linaria nivea
Linaria nurensis
Linaria oblongifolia
Linaria odora
Linaria oligantha
Linaria onubensis
Linaria paradoxa
Linaria parviracemosa
Linaria pedicellata
Linaria pedunculata
Linaria pelisseriana
Linaria peloponnesiaca
Linaria peltieri
Linaria pinifolia
Linaria platycalyx
Linaria polygalifolia
Linaria popovii
Linaria propinqua
Linaria pseudolaxiflora
Linaria pseudoviscosa
Linaria purpurea
Linaria pyramidalis
Linaria ramosa
Linaria reflexa
Linaria remotiflora
Linaria repens
Linaria ricardoi
Linaria riffea
Linaria rubioides
Linaria salzmannii
Linaria satureioides
Linaria saxatilis
Linaria schelkownikowii
Linaria schirvanica
Linaria sessilis
Linaria simplex
Linaria spartea
Linaria striatella
Linaria supina
Linaria tarhunensis
Linaria tenuis
Linaria thibetica
Linaria thymifolia
Linaria tingitana
Linaria tonzigii
Linaria transiliensis
Linaria triornithophora
Linaria triphylla
Linaria tristis
Linaria tuberculata
Linaria tursica
Linaria unaiensis
Linaria venosa
Linaria ventricosa
Linaria veratrifolia
Linaria verticillata
Linaria virgata
Linaria viscosa
Linaria vulgaris
Linaria warionis
Linaria weilleri
Linaria yunnanensis

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