Evolution in SNAPDRAGONS

Linaria - Linaria virgata - Taxonomy

Linaria virgata (Poiret) Desf., Fl. Atl. 2: 41, tab. 135 (1798).

Glabrous annual; fertile stems erect or ascending, branching from base, scarcely above; sterile stems occasionally present, procumbent. Leaves of fertile stems flat, attenuate or cuneate, 3­ or 4-verticillate below, alternate above; leaves of sterile stems similar but usually smaller, 3-verticillate. Inflorescence usually dense in flower, becoming lax in fruit. Bracts 1.5-8 mm, oblong or linear, acute or subacute, deflexed. Pedicels 0.5-1 mm in flower, 1-3 mm in fruit, more or less erect. Calyx-lobes unequal, oblong to lanceolate. Corolla reddish-purple or lilac, occasionally pink or white, with tube marked with yellow; spur slender, straight or somewhat curved forwards. Capsule glabrous. Seeds irregularly tetrahedral, the margins acute or obtuse, without longitudinal ridge, the intervening faces somewhat coarsely ruminate-alveolate or ruminate-foveate, blackish-grey or black; hilum subbasal, narrow; periclinal wall of testa-cells concave with prominent margins, not papillate, verruculate to rugulate. Modified from Sutton (1988).

North Africa. Ag Li Ma? Tn.

Type: ALGERIA: environs de La Calle [El Kala] Poiret s.n. (holo.? (cf. Viano, 1978b: 242) P-LA).



a. Linaria virgata subsp. virgata.

Fertile stems 1-8 (-12), 12-60 cm, suberect or erect, mostly simple, sterile stems 0-7. Leaves of fertile stems 7-19 x 2-8 mm, narrowly oblanceolate to obovate or elliptic, mostly acute. Inflorescence with (7-) 15-40 flowers, dense in flower, dense or somewhat lax in fruit. Calyx­lobes setaceous or linear-lanceolate, long-acuminate; adaxial lobe 3-7 x 0.5-1 mm; abaxial lobes 2.2-5.3 x 0.6-1.4 mm. Corolla 18-24 mm, bluish-violet, the palate lilac, the spur deep violet; tube 1.5-2 mm broad at mouth; adaxial lip sinus 3-4.2 mm; abaxial lip sinus 1.3-2 mm; spur 9-13.5 mm, 0.7-1.1 mm broad at base, equalling or exceeding rest of corolla. Capsule 2.5-4 x 3-4.5 mm, subglobose. Seeds 1.1-1.4 mm.

NE AIgeria & NW Tunisia. Ag Tn.



b. Linaria virgata subsp. tunetana Murb., Lunds Univ. Årrskr. ser. 1 34(2/7): 27, tab. 9(12-13) (1898).

Fertile stems 1-9, 4-15 cm, ascending, simple or branched; sterile stems apparently absent. Leaves of fertile stems 7-18 x 2-10 mm, obovate or ellíptic, obtuse to acuminate. Inflorescence with 2-8 flowers, dense in flower, somewhat lax in fruit. Calyx-lobes linear-oblanceolate or oblong-oblanceolate, obtuse to subacute; adaxial lobe 6.5-8 x 1-1.5 mm; abaxial lobes 4-6 x 1-2 mm. Corolla 21-27 mm, bluish-violet; tube 3-4 mm broad at mouth; adaxial lip sinus 3.5­4.5 mm; abaxial lip sinus 1.5-2.5 mm; spur 9-11 mm, 1.1-1.5 mm broad at base, shorter than rest of corolla. Capsule 4-5 x 4-5 mm, subglobose. Seeds 1.2-1.5 mm.

C. Tunisia. Tn.

Type: TUNISIA: Tunetia media, Maktar, in collib. calcareis, 3 vi 1896 Murbeck s .n. (lectotype, selected here LD, isolecto. BM!).



c. Linaria virgata subsp. algeriensis Murb., Lunds Univ. Årrskr. ser. 1 34(2/7): 25 (1898).

Fertile stems usually numerous, 5-18 (-20) cm, decumbent to ascending, rarely suberect, usually simple; sterile stems usually absent. Leaves of fertile stems 7-28 x (3-) 5-11 mm, oblong, obovate or suborbicular, acute to obtuse. Inflorescence with 3-15 flowers, dense in flower, becoming somewhat lax in fruit. Calyx-lobes linear to oblong, obtuse to acute; adaxial lobe (3.2-) 5-6 x 0.5-0.9 mm; lateral lobes (2.4-) 3.5-4 x 0.7-1.4 mm. Corolla (14-) 20-25 mm, reddish-purple or occasionally white with yellow palate and lilac spur; tube 2.5-3 mm broat at mouth; adaxial lip sinus 2-2.8 mm; abaxial lip sinus 0.9-1.7 mm; spur (7-) 8-15 mm, (0.8-) 1.5-3 mm broad at base, more or less equalling rest of corolla. Capsule 3-4 x 3.5-5 mm, globose. Seeds 1-1.5 mm.

N Africa. Ma? Ag Tn.

Type: ALGERIA: [C1/H2]; Constantine, Kroubs [El Khroub], 11 iii 1896 Murbeck& Olin s.n. (lectotype, selected here, LD, isolecto. BM!).



d. Linaria virgata subsp. syrtica Murb., Lunds Univ. Årrskr. ser. 1 34(2/7): 26 (1898).

Fertile stems 5-30, 5-11 (-20) cm, decumbent to ascending, simple or branched; sterile stems 0 (-5). Leaves 5-16 (-25) x 2-5 mm, oblong to elliptic, acute or subacute. Inflorescence with (2-) 3-14 flowers, capitate or subcapitate in flower, becoming somewhat lax in fruit. Calyx-lobes linear to oblong, obtuse to subacute; adaxíal lobe 3.5-4.5 (-5.5) x 0.7-1 (-1.3) mm; abaxíal lobes 2.5-3.5 (-4.5) x 0.7-1.1 (-2) mm. Corolla 12-17 (-21) mm, bluish-víolet or reddish­purple, the palate lilac or white; tube 2-2.5 mm broad at base, adaxial lip sínus 2.5-3 mm; abaxial líp sínus 1-1.5 mm; spur 5-7 (-9) mm, 0.8-1.4 (-1.6) mm broad at base, shorter than rest of corolla. Capsule 4-5 x 3.5-4.5 mm, oblong-ovoid. Seeds 0.7-1 mm.

CN Africa. Lí Tn.

Type: TUNISIA: Oasis de Métouia, 4 iv 1896 Murbeck s.n. (lectotype, selected here, LD, isolecto. BM!).



Linaria species
Linaria aeruginea
Linaria albifrons
Linaria algarviana
Linaria alpina
Linaria altaica
Linaria amethystea
Linaria amoi
Linaria angustissima
Linaria anticaria
Linaria antilibanotica
Linaria arenaria
Linaria armeniaca
Linaria arvensis
Linaria atlantica
Linaria badalii
Linaria baligaliensis
Linaria bamianica
Linaria biebersteinii
Linaria bipartita
Linaria bipunctata
Linaria bordiana
Linaria brachyceras
Linaria bubanii
Linaria bungei
Linaria buriatica
Linaria caesia
Linaria capraria
Linaria cavanillesii
Linaria chalepensis
Linaria clementei
Linaria confertiflora
Linaria corifolia
Linaria cossonii
Linaria coutinhoi
Linaria cretacea
Linaria creticola
Linaria dalmatica
Linaria damascena
Linaria debilis
Linaria decipiens
Linaria depauperata
Linaria diffusa
Linaria dissita
Linaria elegans
Linaria elymaitica
Linaria fallax
Linaria fastigiata
Linaria faucicola
Linaria ficalhoana
Linaria filicaulis
Linaria flava
Linaria genistifolia
Linaria gharbensis
Linaria glacialis
Linaria glauca
Linaria grandiflora
Linaria griffithsii
Linaria haelava
Linaria hellenica
Linaria hepatica
Linaria hirta
Linaria huteri
Linaria iconia
Linaria incarnata
Linaria incompleta
Linaria intricata
Linaria japonica
Linaria joppensis
Linaria kokanica
Linaria kulabensis
Linaria kurdica
Linaria latifolia
Linaria laxiflora
Linaria leptoceras
Linaria lilacina
Linaria lineolata
Linaria loeselii
Linaria macrophylla
Linaria macroura
Linaria maroccana
Linaria melampyroides
Linaria meyeri
Linaria michauxii
Linaria micrantha
Linaria microsepala
Linaria miller
Linaria multicaulis
Linaria munbyana
Linaria nivea
Linaria nurensis
Linaria oblongifolia
Linaria odora
Linaria oligantha
Linaria onubensis
Linaria paradoxa
Linaria parviracemosa
Linaria pedicellata
Linaria pedunculata
Linaria pelisseriana
Linaria peloponnesiaca
Linaria peltieri
Linaria pinifolia
Linaria platycalyx
Linaria polygalifolia
Linaria popovii
Linaria propinqua
Linaria pseudolaxiflora
Linaria pseudoviscosa
Linaria purpurea
Linaria pyramidalis
Linaria ramosa
Linaria reflexa
Linaria remotiflora
Linaria repens
Linaria ricardoi
Linaria riffea
Linaria rubioides
Linaria salzmannii
Linaria satureioides
Linaria saxatilis
Linaria schelkownikowii
Linaria schirvanica
Linaria sessilis
Linaria simplex
Linaria spartea
Linaria striatella
Linaria supina
Linaria tarhunensis
Linaria tenuis
Linaria thibetica
Linaria thymifolia
Linaria tingitana
Linaria tonzigii
Linaria transiliensis
Linaria triornithophora
Linaria triphylla
Linaria tristis
Linaria tuberculata
Linaria tursica
Linaria unaiensis
Linaria venosa
Linaria ventricosa
Linaria veratrifolia
Linaria verticillata
Linaria virgata
Linaria viscosa
Linaria vulgaris
Linaria warionis
Linaria weilleri
Linaria yunnanensis

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