Evolution in SNAPDRAGONS

Linaria - Linaria viscosa - Taxonomy

Linaria viscosa (L.) Chaz., Suppl. Dict. Jard. 2: 39 (1790).

Annual; fertile stems simple or branched above; sterile stems usually several. Leaves of fertile stems 5-32 x (0.3-) 0.5-2 mm, linear, obtuse, flat, alternate above, 4-verticillate below; leaves of sterile stems elliptic, acute or acuminate, 3- or 4-verticillate. Inflorescence with 1-20 flowers, dense in flower and fruit. Bracts 2-6 mm. Pedicels erect. Calyx-lobes linear-lanceolate, acute, margin scarious. Corolla bright yellow; spur straight, somewhat shorter than rest of corolla. Style deeply bifid. Capsule 3.5-5 x 3-4 mm, oblong-ovoid, glabrous; loculi equal or subequal, each loculus dehiscing by 3 teeth. Seeds 0.5-0.7 mm, blackish-grey; periclinal wall of testa-cells verrucate or verruculate, the margin prominently raised, forming rounded marginal papilla towards ridge-apex, the median papilla usually present, small, rounded, more or less isodiametric. Modified from Sutton (1988).

S & W Iberian peninsula, NW Africa? Hs Lu Ma?

Type: SPAIN: "viscosum" (syn. (cf. Viano, 1978a: 57) LINN-767.24!);  "viscosum" (syn. (cf. Viano, 1978a: 57) LINN-767.25!).




a. Linaria viscosa subsp. viscosa.

Glabrous except for villous inflorescence; fertile stems 1-6, 17-75 cm, erect or sometimes ascending. Leaves of sterile stems elliptic, acute, 3- or 4-verticillate. Inflorescence more or less dense in flower and fruit. Bracts linear to elliptic, acute. Pedicels free, 5-10 mm in flower, up to 12 mm in fruít. Calyx-lobes 3.8-7 x 0.9-1.6 mm, linear-lanceolate. Corolla 24-26 mm; adaxial lip sinus 3-4 mm; abaxial lip sinus 0.6-0.9 mm; spur 9-10 mm, 1-1.5 (-2) mm broad. Seeds strongly transversely ridged; ridges 5-8, usually discrete, rounded to subobtuse.

2n = 12 (Valdés, 1970b; Löve and Kjellqvist, 1974: 184).

S Spain, S Portugal, N Morocco? Hs Lu Ma?


b. Linaria viscosa subsp. crassifolia (Coutinho) D.A. Sutton, stat. nov.

Basionym: Linaria viscosa var. crassifolia Coutinho, Fl. Port.: 550 (1913).

Glabrous except for somewhat sparsely villous inflorescence; fertile stems 2-10, 9-26 cm, decumbent or arcuate-ascending. Leaves of sterile stems linear to elliptic, obtuse, 4-verticillate, fleshy. Inflorescence dense and often subcapitate in flower, dense to more or less lax in fruit. Bracts elliptic to ovate, acute. Pedicels free; 4-10 mm in flower, 9-17 (-24) mm in fruit. Calyx­lobes 3.7-5.5 x 0.9-1.7 mm, linear-lanceolate. Corolla 21-28 mm, bright yellow, often with the adaxial lip purple in bud; adaxial lip sinus 2.5-3 mm; abaxial lip sinus 1-1.2 mm; spur 9-11 mm, 1.5-2.5 broad. Seeds weakly transversely ridged or ruminate-alveolate; ridges 4-6, often anastomosed, rounded.

CW Portugal. Lu.

Type: PORTUGAL: Estremadura; prox. Cabo da Roca (not located).



c. Linaria viscosa subsp. spicata (Coutinho) D.A. Sutton, comb. et stat. nov.

Basionym: Linaria spicata G. Kunze, Flora (Regensb.) 29(41): 645 (1846).

Glabrous except for glandular-pubescent inflorescence and base of fertile stems; fertile stems 1-12, 8-40 cm, ascending to erect, slender. Leaves of sterile stems linear to elliptic, obtuse, 3- or 4- verticillate. Inflorescence dense in flower and fruit. Bracts linear to oblong, obtuse. Pedicels 2-6 mm in flower, up to 10 mm in fruit, occasionally adnate for 0.25-0.3 of length to stem, erect. Calyx-lobes 2.8-4.5 x 0.4-0.9 mm, linear. Corolla 14-20 mm, yellow; adaxial lip sinus 1-2 mm; abaxial lip sinus 0.5-0.8 mm; spur 6.5-9 mm; 0.6-1 mm broad. Seeds transversely ridged or ruminate-alveolate; ridges 6-11, discrete or anastomosed, rounded to subacute.

S Spain (mainly Sierra Nevada). Hs.

Type: SPAIN: [Granada]; Sierra Nevada, "in latere australi ad rivulos regionis alpinae", ix 1844 Willkomm s.n. Herb. Kunze (syn.? LZ†); Sierra Nevada, 6000-7000 ft, viii 1844 Willkomm s.n. (type collection? BM!).










Linaria species
Linaria aeruginea
Linaria albifrons
Linaria algarviana
Linaria alpina
Linaria altaica
Linaria amethystea
Linaria amoi
Linaria angustissima
Linaria anticaria
Linaria antilibanotica
Linaria arenaria
Linaria armeniaca
Linaria arvensis
Linaria atlantica
Linaria badalii
Linaria baligaliensis
Linaria bamianica
Linaria biebersteinii
Linaria bipartita
Linaria bipunctata
Linaria bordiana
Linaria brachyceras
Linaria bubanii
Linaria bungei
Linaria buriatica
Linaria caesia
Linaria capraria
Linaria cavanillesii
Linaria chalepensis
Linaria clementei
Linaria confertiflora
Linaria corifolia
Linaria cossonii
Linaria coutinhoi
Linaria cretacea
Linaria creticola
Linaria dalmatica
Linaria damascena
Linaria debilis
Linaria decipiens
Linaria depauperata
Linaria diffusa
Linaria dissita
Linaria elegans
Linaria elymaitica
Linaria fallax
Linaria fastigiata
Linaria faucicola
Linaria ficalhoana
Linaria filicaulis
Linaria flava
Linaria genistifolia
Linaria gharbensis
Linaria glacialis
Linaria glauca
Linaria grandiflora
Linaria griffithsii
Linaria haelava
Linaria hellenica
Linaria hepatica
Linaria hirta
Linaria huteri
Linaria iconia
Linaria incarnata
Linaria incompleta
Linaria intricata
Linaria japonica
Linaria joppensis
Linaria kokanica
Linaria kulabensis
Linaria kurdica
Linaria latifolia
Linaria laxiflora
Linaria leptoceras
Linaria lilacina
Linaria lineolata
Linaria loeselii
Linaria macrophylla
Linaria macroura
Linaria maroccana
Linaria melampyroides
Linaria meyeri
Linaria michauxii
Linaria micrantha
Linaria microsepala
Linaria miller
Linaria multicaulis
Linaria munbyana
Linaria nivea
Linaria nurensis
Linaria oblongifolia
Linaria odora
Linaria oligantha
Linaria onubensis
Linaria paradoxa
Linaria parviracemosa
Linaria pedicellata
Linaria pedunculata
Linaria pelisseriana
Linaria peloponnesiaca
Linaria peltieri
Linaria pinifolia
Linaria platycalyx
Linaria polygalifolia
Linaria popovii
Linaria propinqua
Linaria pseudolaxiflora
Linaria pseudoviscosa
Linaria purpurea
Linaria pyramidalis
Linaria ramosa
Linaria reflexa
Linaria remotiflora
Linaria repens
Linaria ricardoi
Linaria riffea
Linaria rubioides
Linaria salzmannii
Linaria satureioides
Linaria saxatilis
Linaria schelkownikowii
Linaria schirvanica
Linaria sessilis
Linaria simplex
Linaria spartea
Linaria striatella
Linaria supina
Linaria tarhunensis
Linaria tenuis
Linaria thibetica
Linaria thymifolia
Linaria tingitana
Linaria tonzigii
Linaria transiliensis
Linaria triornithophora
Linaria triphylla
Linaria tristis
Linaria tuberculata
Linaria tursica
Linaria unaiensis
Linaria venosa
Linaria ventricosa
Linaria veratrifolia
Linaria verticillata
Linaria virgata
Linaria viscosa
Linaria vulgaris
Linaria warionis
Linaria weilleri
Linaria yunnanensis

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