Evolution in SNAPDRAGONS


Holzneria - Taxonomy



Holzneria Speta, Bot. Jahrb. 103(1): 16 (1982). Annual herbs; stems homomorphic, erect, the branches not cirrhous. Leaves homomorphic, simple, entire, pinnately veined, petiolate, opposite below, alternate above; petioles not cir- rhous. Flowers zygomorphic, shortly pedicellate or subsessile, in lax, leafy, spike-like racemes. Floral leaves not subtending additional axillary shoots. Pedicels not cirrhous. Calyx deeply divided, the lobes entire, valvate, conspicuously unequal, the adaxial lobe much shorter than lateral lobes, the lateral lobes exceeding corolla-tube. Corolla with tube cylindrical, calcarate abaxially at base; limb bilabiate, the lips subequal, the adaxial lip shallowly emarginate, erecto- patent, the abaxial lip spreading with low palate more or less occluding mouth of tube, the spur conical, straight, inconspicuous; lobes entire. Fertile stamens 4, didynamous, included, the adjacent pairs of anthers marginally coherent, the connective not dilated above anther; staminode minute. Style simple, erect; stigma capitate, entire, positioned between anthers of fertile stamens. Capsule ovoid to subglobose, often with persistent corolla, the walls somewhat thick and coriaceous; loculi unequal, the adaxial loculus exceeding abaxial, many-seeded, dehiscing from apex by 4-6 irregular valves forming solitary pore, the abaxial loculus few- seeded, indehiscent or dehiscing after abaxial by irregular valves forming solitary pore. Seeds more or less radially symmetrical or somewhat dorsi-ventrally asymmetrical, ovoid to narrowly ovoid, truncate, reticulate-alveolate; hilum more or less basal; ridges regularly anastomosed; periclinal wall of testa-cells tabular or tabular-convex, that of cells from interstices with median papilla; epicuticular waxes sometimes present, granulate. By Sutton (1988)

Holzneria species
H. microcentron
H. spicata

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